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For the relationships of Boraginales, see discussion under Gentianales.īORAGINALES Berchtold & J. For the complex patterns of variation in a number of characters in this part of the tree, see the Gentianales page. Nectary at base of G style long several ovules/carpel fruitĮvolution: Divergence & Distribution. : (herbaceous habit widespread) (8-ring deoxyflavonols +) vesselĮlements with simple perforation plates nodes 1:1 C forming a distinct tube, initiation late A epipetalous (vascularized) ]: G, superposed loss of introns 18-23 in RPB2 gene d copy. Our online platform, Wiley Online Library () is one of the world’s most extensive multidisciplinary collections of online resources, covering life, health, social and physical sciences, and humanities.Gametophyte dominant, independent, multicellular, initially ±globular, not motile, branched showing gravitropism glycolate oxidase +, glycolate metabolism in leaf peroxisomes, acquisition of phenylalanine lysase*, flavonoid synthesis*, microbial terpene synthase-like genes +, triterpenoids produced by CYP716 enzymes, CYP73 and phenylpropanoid metabolism, xyloglucans in primary cell wall, side chains charged plant poikilohydrous, ectohydrous thalloid, leafy, with single-celled apical meristem, tissues little differentiated, rhizoids +, unicellular chloroplasts several per cell, pyrenoids 0 centrioles/centrosomes in vegetative cells 0, microtubules with γ-tubulin along their lengths, interphase microtubules form hoop-like system metaphase spindle anastral, predictive preprophase band +, phragmoplast +, plasmodesmata + antheridia and archegonia +, jacketed*, surficial blepharoplast +, centrioles develop de novo, bicentriole pair coaxial, separate at midpoint, centrioles rotate, associated with basal bodies of cilia, multilayered structure + (0), spline +, basal body 200-250 nm long, associated with amorphous electron-dense material, microtubules in basal end lacking symmetry, stellate array of filaments in transition zone extended, axonemal cap 0 male gametes with a left-handed coil, cilia 2, lateral, asymmetrical oogamy sporophyte +*, multicellular, growth 3-dimensional*, cuticle +*, plane of first cell division transverse, sporangium and upper part of seta developing from epibasal cell, with at least transient apical cell, initially surrounded by and dependent on gametophyte, placental transfer cells +, in both sporophyte and gametophyte, wall ingrowths develop early suspensor/foot +, cells at foot tip somewhat haustorial sporangium +, single, terminal, dehiscence longitudinal meiosis sporic, monoplastidic, MTOC associated with plastid, sporocytes 4-lobed, cytokinesis simultaneous, preceding nuclear division, quadripolar microtubule system + wall development both centripetal and centrifugal, 1000 spores/sporangium, sporopollenin in the spore wall* laid down in association with trilamellar layers plastid transmission maternal nuclear genome seed single duplication of the PI gene. With a growing open access offering, Wiley is committed to the widest possible dissemination of and access to the content we publish and supports all sustainable models of access. Wiley has partnerships with many of the world’s leading societies and publishes over 1,500 peer-reviewed journals and 1,500+ new books annually in print and online, as well as databases, major reference works and laboratory protocols in STMS subjects.
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